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Authors
Shito, Takumi T.
Jayakumar, Vasanthan https://orcid.org/0000-0002-6067-4184
Nishitsuji, Koki
Nishitsuji, Yoshie
Kawai, Shimon
Miyasaka, Shunsuke O.
Oka, Kotaro
Sakakibara, Yasubumi
Hotta, Kohji https://orcid.org/0000-0003-4614-7473
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Funding
Funding for this research was provided by:
The Research Institute of Marine Invertebrates (IKU2021-02)
The Keio University Doctorate Student Grant-in-Aid Program from the Ushioda Memorial Fund
JSPS KAKENHI (JP 22J22628 and 25KJ0268)
JSPS KAKENHI (21H00440, 23H04717, 24K02038, and 25H01799)
the Keio University Research and Education Center for Natural Sciences Budget
the KLL Keio Leading Program
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Article History
Received: 6 November 2025
Accepted: 23 June 2026
First Online: 4 August 2026
Declarations
Ethics approval and consent to participate: Not applicable.
Consent for publication: Not applicable.
Competing interests: The authors declare no competing interests.
