Fan, Saibin
Lu, Yang
Yue, Caipeng
Zhu, Shixin
Huang, Jinyong
Wang, Yuhui
Lai, Yong
Zhang, Ningning
Funding for this research was provided by:
National Natural Science Foundation of China (31800551)
National Natural Science Foundation of China (81903747)
Science and Technology Research Project of Henan Province (252102110244)
Article History
Received: 17 November 2025
Accepted: 27 April 2026
First Online: 1 May 2026
Declarations
:
: The authors confirm that permission for the collection of plant materials from all mountainous field sites was obtained from the relevant local forestry and natural resources authorities. The field studies complied with relevant institutional, national, and international guidelines and legislation. The voucher specimens (voucher numbers: LY2019063003, LY2020061434, LY2020071509, LY2023051709, LY2023052101, LY2023062601, LY2023062701, LY2023062706) have been deposited in the Herbarium of Zhengzhou University and were identified by Yang Lu from Zhengzhou University.
: Not applicable.
: The authors declare no competing interests.