La, Boya
Jiang, Chunyan
He, Jia
Ni, Man
Zhou, Jing
Wu, Huimin
Ning, Song
Dai, Wangjuan
Yan, Zhengjie
Cui, Yugui
Qin, Lianju
Liu, Jiayin
Funding for this research was provided by:
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
National Natural Science Foundation of China (81730041)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
State Key Laboratory of Reproductive Medicine Program (SKLRM-GC201803)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
National Key Research and Development Program of China (2021YFC2700404)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
the National Key Research and Development Program of China (2021YFC2700501)
Article History
Received: 18 July 2025
Accepted: 22 November 2025
First Online: 2 January 2026
Change Date: 26 March 2026
Change Type: Update
Change Details: The original online version of this article was revised: the funding section has been corrected.
Declarations
:
: All animal studies (including mouse euthanasia procedures) were performed in compliance with the regulations and guidelines of Nanjing Medical University institutional animal care (Approval number: IACUC-2004028) and conducted according to the AAALAC and IACUC guidelines. All methods are reported in accordance with ARRIVE guidelines.
: Not applicable.
: The authors declare no competing interests.