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Authors
Toyoshi, Eita
Watanabe, Masahiro
Namai, Fu
Yamane, Kenji
Kashima, Toma
Ikeda-Ohtsubo, Wakako
Agista, Afifah Zahra
Yoshida, Ayu
Sakuma, Taiga
Fukuda, Itsuko
Suzuki, Kasumi
Yoshioka, Gou
Imai, Yuji
Tsuchida, Sae
Nishiyama, Eri
Shinkai, Hiroki
Muneta, Yoshihiro
Uenishi, Hirohide
Tobe, Ryuta
Shirakawa, Hitoshi
Maekawa, Masamitsu
Mano, Nariyasu
Kitazawa, Haruki
Nishiyama, Keita
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Funding
Funding for this research was provided by:
Japan Society for the Promotion of Science (Grant-in-Aid for Challenging Research (Exploratory) (23K18072))
Japan Society for the Promotion of Science (Grant-in-Aid for Scientific Research (B) (23K27051))
Bio-oriented Technology Research Advancement Institution (Development of Innovative Technology grant (JPJ007097))
Japan Racing Association
Uehara Memorial Foundation
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Article History
Received: 18 September 2025
Accepted: 21 January 2026
First Online: 18 February 2026
Declarations
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: The study protocol was approved by the Animal Care and Management Committee (approval numbers: Miyagi Pref. Livestock Experiment Station R3-2-2(4) and No. 2016-23). All procedures for sample collection and processing adhered to guidelines set by the Biosafety Committee of Tohoku University (approval number: 2016-045-01). Sampling was conducted in accordance with standard animal welfare and husbandry regulations at each participating farm.
: Not applicable.
: Author Kenji Yamane is employees of Nihon Shokuhin Kako Co. Ltd. Authors Sae Tsuchida and Eri Nishiyama is employees of FASMAC Co., Ltd. The remaining authors declare no competing interests.
