Shi, Fuyu
Yang, Zhiteng
Zhang, Liangzhi
Zou, Desheng
Yu, Jiangkun
Guo, Na
Ren, Shien
Tang, Xianjiang
Gu, Chen
Xu, Ruiping
Ru, Yuning
Zhang, Yanming
Wang, Dehua
Funding for this research was provided by:
National Natural Science Foundation of China (32301301)
National Natural Science Foundation of China (32330012)
Natural Science Foundation of Qingdao (23-2-1-26-zyyd-jch)
Natural Science Foundation of Shandong (ZR2024QC355)
Double-First Class Initiative of the School Life Sciences, Shandong University
Article History
Received: 6 November 2024
Accepted: 14 August 2025
First Online: 12 December 2025
Ethics approval and consent to participate
: This work was approved by the Animal Care and Use Committee of Shandong University for animal experimentation and research. Samples were collected with informed consent from all participants.
: Not applicable.
: The authors declare no competing interests.