Funding for this research was provided by:
Guangdong South China Sea Key Laboratory of Aquaculture for Aquatic Economic Animals (KFKT2019ZD04)
the Open Fund of Shandong Key Laboratory of Disease Control in Mariculture (KF201802)
the Shanghai Pujiang Talent Program (18PJ1404000)
Shanghai Municipal Natural Science Foundation (17ZR1412900)
Natural Science Foundation of China (31872587)
Received: 10 October 2019
Accepted: 7 January 2020
First Online: 24 January 2020
Ethics approval and consent to participate
: All applicable international, national, and/or institutional guidelines for the care and use of animals were followed.
: Not applicable.
: The authors declare that they have no competing interests.