Funding for this research was provided by:
Chinese Academy of Sciences (KZCX2-YW-Q07-04)
Natural Science Foundation of Shandong Province (ZR2018MD012)
National Natural Science Foundation of China (42072326, 41772348, 40901027)
Received: 21 April 2021
Accepted: 19 November 2021
First Online: 2 December 2021
: This study does not involve human participants, animals followed all applicable international and national guidelines for use of animals in research. All authors have consented to participate.
: Not applicable.
: The authors declare that they have no competing interests.