Funding for this research was provided by:
Horizon 2020 Framework Programme (679302)
Biotechnology and Biological Sciences Research Council (BBS/E/D/30002275)
Rural and Environment Science and Analytical Services Division (Not applicable)
Received: 24 June 2019
Accepted: 29 October 2019
First Online: 12 November 2019
Ethics approval and consent to participate
: All authors declare that animal samples were obtained in compliance with local/national laws in force at the time of sampling. Data exchange was in accordance with national and international regulations, and approved by the owners.
: All authors have read and approved the final manuscript.
: The authors declare that they have no competing interests.