Funding for this research was provided by:
National Natural Science Foundation of China (31670496)
Fundamental Research Funds for the Central Universities (2572015EA02)
Received: 23 November 2018
Accepted: 28 August 2019
First Online: 13 September 2019
Ethics approval and consent to participate
: Sampling permission has been obtained for Changbai Mountain National Nature Reserve in Jilin Province, Northeastern China and field studies were conducted in accordance with local legislation.
: Not applicable.
: The authors declare that they have no competing interests.