Funding for this research was provided by:
the Joint Fund of the National Natural Science Foundation of China and the Karst Science Research Center of Guizhou Province (U1812401)
Provincial Program on Platform and Talent Development of the Department of Science and Technology of Guizhou China (2019-5661, 2019-5617)
Received: 27 March 2020
Accepted: 28 April 2020
First Online: 29 May 2020
Ethics approval and consent to participate
: The permission to enter the study sites and to collect the samples was obtained from the Guizhou Zhijin cave Global Geopark, China.
: Not applicable.
: The authors declare that they have no competing interests.