Funding for this research was provided by:
National Natural Science Foundation of China (31471352, 31501953, 31501161, 31501198)
Received: 15 June 2018
Accepted: 20 September 2018
First Online: 3 October 2018
: All animals were kept in accordance with the protocols approved by the guidelines of the Institutional Animal Care and Use Committee of the Institute of Zoology (IOZ), Chinese Academy of Sciences (CAS), Beijing, China.
: Not applicable.
: The authors declare that they have no competing interests.
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