Funding for this research was provided by:
The Darwin Trust
Wellcome Trust (202769/Z/16/Z)
Royal Society (UF110155)
Received: 31 October 2019
Accepted: 7 January 2020
First Online: 14 January 2020
Ethics approval and consent to participate
: All procedures were carried out in accordance with the UK Home Office regulations (Animals Scientifc Procedures Act 1986; project licence number: 70/8546) and approved by the University of Edinburgh.
: Not applicable.
: The authors declare that they have no competing interests.