Funding for this research was provided by:
National Natural Science Foundation of China (81572014)
Pearl River Nova Program of Guangzhou (201710010030)
National High Technology Research and Development Program of China (2015AA020934)
National Research and Development Plan of China (2016YFC1200500)
Doctoral Program of Higher Education of China (20120171120049)
National Science Foundation of Guangdong Province (S2012040007256)
Received: 8 November 2017
Accepted: 18 January 2018
First Online: 2 February 2018
: All the animal experiments were approved by the medical research ethics committee of Sun Yat-sen University and conformed to the Chinese National Institute of Health Guide for the Care and Use of Laboratory Animals (No. 2017-020).
: Not applicable
: The authors declare that they have no competing interests.
: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.