Funding for this research was provided by:
National Natural Science Foundation of China (31672306, 31572277)
Strategic Priority Research Program of the Chinese Academy of Sciences (XDB11010400)
Foundation of State Key Laboratory of IPM (ChineseIPM1701)
Received: 8 August 2017
Accepted: 2 February 2018
First Online: 23 February 2018
: The procedures for animal care and use were performed in strict accordance with the guidelines of the Animal Use Committee of the Institute of Zoology, Chinese Academy of Sciences (IOZ 2015).
: Not applicable.
: The authors declare that they have no competing interests.
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