Funding for this research was provided by:
National Natural Science Foundation of China (31872478)
National Key Research and Development Programs of China (2016YFD0500103)
Natural Science Foundation of Shanghai (19ZR1469100)
China Postdoctoral Science Foundation (2017M611074)
Received: 7 December 2019
Accepted: 26 February 2020
First Online: 3 April 2020
Ethics approval and consent to participate
: The study was approved by the Ethics and Animal Welfare Committee of Shanghai Veterinary Research Institute, China and the animals were handled by the rules stipulated by the Ethics and Animal Welfare Committee of Shanghai Veterinary Research Institute, China.
: Not applicable.
: The authors declare that they have no competing interests.