Funding for this research was provided by:
Biotechnology and Biological Sciences Research Council (BB/J004669/1, BB/J010375/1, BB/CSP17270/1, BB/M011216/1)
Received: 31 July 2019
Accepted: 27 September 2019
First Online: 10 October 2019
Ethics approval and consent to participate
: Not applicable.
: This study abides by UK guidelines and legislation for plant science research.
: The authors declare that they have no competing interests.