Funding for this research was provided by:
Japan Agency for Medical Research and Development (JP18DM0107056)
Japan Society for the Promotion of Science (16K15478)
Ministry of Education, Culture, Sports, Science and Technology (10036021)
Received: 16 October 2018
Accepted: 15 March 2019
First Online: 12 April 2019
Ethics approval and consent to participate
: All animal experimental procedures were performed in accordance with the regulations of the University of Tokyo and were approved by the committee of the University of Tokyo Graduate School of Medicine.
: Not applicable.
: The authors declare that they have no competing interests.
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