Funding for this research was provided by:
National Natural Science Foundation of China (31711530221, 31501011, 91731303, 31771388)
National Science Fund for Distinguished Young Scholars (31525014)
Program of Shanghai Academic Research Leader (16XD1404700)
National Key Research and Development Program of China (2016YFC0906403)
Strategic Priority Research Program (XDB13040100)
Key Research Program of Frontier Sciences (QYZDJ-SSW-SYS009)
UK Royal Society-Newton Advanced Fellowship (NAF\R1\191094)
Shanghai Municipal Science and Technology Major Project (2017SHZDZX01)
Chinese Academy of Sciences President’s International Fellowship Initiatives (2017VBA0008)
Received: 25 April 2019
Accepted: 26 September 2019
First Online: 22 October 2019
Ethics approval and consent to participate
: All samples were collected with informed consent and approved by the Biomedical Research Ethics Committee of the Shanghai Institutes for Biological Sciences (No. ER-SIBS-261408).
: Not applicable for public available data collected from previous publications. All participants donated DNA for this study have provided consent for publication of summary statistics on the population level.
: The authors declare that they have no competing interests.