Funding for this research was provided by:
Biotechnology and Biological Sciences Research Council (BB/K014773/1, BB/M011208/1, BB/P01738X/1)
Received: 12 July 2019
Accepted: 5 October 2019
First Online: 15 October 2019
Ethics approval and consent to participate
: Not applicable.
: All authors consent to publication.
: The authors declare that they have no competing interests.