Funding for this research was provided by:
National Natural Science Foundation of China (31870785, 31570040)
National Key R&D Program of China (No. 2018YFB1501701)
Key Technologies R&D Program of Shandong Province (No. 2018GSF121021)
the Tianjin Municipal Science Foundation (Nos. 18JCQNJC10000)
the State Key Laboratory of Microbial Technology Open Projects Fund
Received: 14 May 2019
Accepted: 9 October 2019
First Online: 15 October 2019
Ethics approval and consent to participate
: Not applicable.
: All authors agreed to the publication of this article.
: The authors declare that they have no competing interests.