Funding for this research was provided by:
Natural Science Foundation of Shandong Province (ZR2018BC014)
National Key Technology R&D Program of China (2015BAD15B05)
National Natural Science Foundation of China (31671985)
Key Technology Research and Development Program of Shandong (2019GNC106060)
Funds of “Taishan Scholar” Construction Project (TS201712023)
Received: 31 August 2019
Accepted: 26 January 2020
First Online: 26 February 2020
Ethics approval and consent to participate
: Not applicable.
: All authors provide their consent for publication of the manuscript in <i>Biotechnology for Biofuels</i>.
: The authors declare that they have no competing interests.