Received: 21 June 2017
Accepted: 29 January 2018
First Online: 20 February 2018
: Animals were maintained in accordance with procedures of The University of Georgia Institutional Animal Care and Use Committee (IACUC), UGA Animal Welfare Assurance #A3437-01, AUP # A2013 06-018-A1.
: Not applicable.
: NCH has conducted research funded by over a dozen pharmaceutical firms; FJ, ES, FS and AF-S are employees of Merck Animal Health.
: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.