Funding for this research was provided by:
National Natural Science Foundation of China (31972707)
national key research and development program of China (2017YFD0501305)
Major scientific research projects (characteristic innovation) of Guangdong Province (2017KTSCX018)
the Guangzhou Zoo Project (h2016141)
the Natural Science Foundation of Guangdong Province (2016A030313396, 2019A1515011534)
Huizhou science and technology projects (2015B040009001)
the Key Research and Development Programme of Guangdong Province (2019B020218004)
Received: 21 March 2019
Accepted: 30 October 2019
First Online: 8 November 2019
Ethics approval and consent to participate
: This study was carried out in accordance with the principles of the Basel Declaration and recommendations of the People’s Republic of China, the Animal Ethics Committee of South China Agricultural University. The protocol was approved by the South China Agricultural University Experimental Animal Welfare Ethics Committee (Approval No. SCXK 2013-0002).
: Not applicable.
: The authors declare that they have no competing interests.