Funding for this research was provided by:
Houghton Trust (Small project research grant)
North of England Zoological Society (Research support funding)
Received: 27 November 2020
Accepted: 21 April 2021
First Online: 21 May 2021
: This project was approved by the science committees of Chester Zoo and Flamingo Land, and by the University of Liverpool Veterinary Research Ethics Committee (reference VREC532a).
: Not applicable.
: The authors declare that they have no competing interests.