Funding for this research was provided by:
Arthritis Australia (1757679)
National Health and Medical Research Council (1043147, 1043147, Senior Principal Research Fellowship)
Deutsche Forschungsgemeinschaft (IRTG 2168)
Received: 6 January 2018
Accepted: 6 March 2018
First Online: 5 April 2018
: Experiments were approved by The University of Melbourne Animal Ethics Committee.
: Not applicable.
: The authors declare that they have no competing interests.
: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.