Funding for this research was provided by:
Arthritis Research UK (20843, 19859)
Wellcome Trust (GGL-104612/Z/14/Z)
Received: 17 December 2018
Accepted: 7 August 2019
First Online: 16 August 2019
Ethics approval and consent to participate
: Experiments were carried out at the University of Birmingham, UK (project licence number P51102987) following strict guidelines governed by the UK Animal (Scientific Procedures) Act 1986 and were approved by the local ethics committee (BERSC: Birmingham Ethical Review Subcommittee).
: Not applicable.
: The authors declare that they have no competing interests.