Funding for this research was provided by:
Japan Society for the Promotion of Science (21370007, 15H04419)
Received: 19 September 2019
Accepted: 16 January 2020
First Online: 30 January 2020
Ethics approval and consent to participate
: All of animal care and breeding were performed according to the guidelines of the animal care and use committee of Tohoku University. The protocol was approved by the committee (permit number: 2017LsA-025).
: Not applicable.
: The authors declare that they have no competing interests.