Funding for this research was provided by:
Biotechnology and Biological Sciences Research Council (BBS/E/D/20002173, BBS/E/D/20002174)
Received: 2 July 2019
Accepted: 11 September 2019
First Online: 22 October 2019
Ethics approval and consent to participate
: The experimental protocol was approved by the Iowa State University Institutional Animal Care and Use Committee (Approval Number 3-17-8475-S) and the Iowa State University Institutional Biosafety Committee (Approval Number 17-I-0011-A). Environmental enrichment of pens was utilized and all pigs were under independent veterinary supervision.
: The authors declare they have no competing interests.