Funding for this research was provided by:
Rufford Foundation (ID0EVSAE252)
Received: 17 July 2018
Accepted: 28 November 2018
First Online: 22 January 2019
Ethics approval and consent to participate
: Ugyen Wangchuck Institute for Conservation and Environmental Research approved this study vide research clearance no UWICE/ADM27/2015-2016/275 dated 07/10/2016.
: Not applicable
: The authors declare that they have no competing interest.
: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.