Funding for this research was provided by:
the Fund of Southern Marine Science and Engineering Guangdong Laboratory (ZJW-2019-07)
National Natural Science Foundation of China (81541153)
Guangdong Provincial Science and Technology Department (2016A050503046, 2015A050502048)
The Public Service Platform of South China Sea for R&D Marine Biomedicine Resources (GDMUK201808)
Received: 17 July 2019
Accepted: 12 November 2019
First Online: 19 November 2019
Ethics approval and consent to participate
: Not applicable.
: All authors consent for publication.
: The authors declare that they have no competing interests.