Funding for this research was provided by:
National Institute of Arthritis and Musculoskeletal and Skin Diseases (R01-AR066663, R00-AR060873, P30-AR057217)
National Institute of Nursing Research (R01-NR015639)
National Human Genome Research Institute (T32 HG000046)
National Institutes of Health (AI074878, AI061570, AI102942, AI095466, AI087990, AI097333, AI095608, AI083480)
Received: 12 June 2017
Accepted: 18 January 2018
First Online: 30 January 2018
Ethics approval and consent to participate
: All mouse experiments were approved by the University of Pennsylvania Institutional Animal Care and Use Committee (IACUC).
: Not applicable.
: The authors declare that they have no competing interests.
: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.