Funding for this research was provided by:
UK Medical Research Council (MC_U105184291)
Received: 15 September 2021
Accepted: 8 November 2021
First Online: 24 November 2021
Change Date: 14 December 2021
Change Type: Update
Change Details: This article has been updated to correct a minor error in the equal contribution statement.
: Experiments were conducted in accordance with the UK Animals (Scientific Procedures) Act of 1986, with local ethical approval (MRC Laboratory of Molecular Biology Animal Welfare and Ethical Review Body Committee). The studies carried out at the UCL Queen Square Institute of Neurology and the Edinburgh Brain Bank were approved through the ethical review processes at each institution.
: The authors declare that they have no competing interest.