Funding for this research was provided by:
National Natural Science Foundation of China (31672296, 31172104)
Received: 9 January 2020
Accepted: 8 May 2020
First Online: 18 May 2020
Ethics approval and consent to participate
: This research complies with the current laws of China, and was approved under the Ethics Committee of School of Life Sciences, Lanzhou University. Field research was conducted with permission from the Bureau of the Gansu An’xi Extremely-Arid Desert National Nature Reserve.
: Not applicable.
: The authors declare that they have no competing interests.