Received: 29 August 2021
Accepted: 25 October 2021
First Online: 8 December 2021
: Our work was performed in accordance with the guidelines for proper conduct of animal experiments in Japan and was approved by the Committee for Animal Experiments in Field Science Center for Northern Biosphere (FSC) of Hokkaido University (ID2–5).
: Not applicable.
: The authors declare that they have no competing interests.