Funding for this research was provided by:
National Geographic Society (WW-079ER-17)
Received: 7 April 2021
Accepted: 7 December 2021
First Online: 20 December 2021
: All research involving animal capture, handling and sample collection was carried out under the Institutional Animal Use and Care Committee of James Madison University (A15-15) and was completed with permission from the University of Brunei Darussalam (UBD/AVC-RI/1.21.1[a]).
: Not applicable.
: The authors declare that they have no competing interests.