Funding for this research was provided by:
National Natural Science Foundation of China (31772443)
National Natural Science Foundation of China (31872214)
National Natural Science Foundation of China (31970387)
the Natural Science Foundation of Fujian Province (2019J05010)
the 111 Project sponsored by the State Bureau of Foreign Experts and Ministry of Education of China (BP2018017)
Article History
Received: 2 December 2019
Revised: 3 April 2020
Accepted: 16 April 2020
First Online: 12 May 2020
Change Date: 20 May 2020
Change Type: Update
Change Details: The copyright line of this article was changed on 20 May 2020 after original online publication.
Change Date: 1 September 2021
Change Type: Erratum
Change Details: An Erratum to this paper has been published:
Change Details: https://doi.org/10.15252/embj.2021109070
Conflict of interest
: The authors declare that they have no conflict of interest.