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Joiner, Aaron MN https://orcid.org/0000-0001-9049-2384

Phillips, Ben P https://orcid.org/0000-0001-9117-4695

Yugandhar, Kumar https://orcid.org/0000-0003-1141-4439

Sanford, Ethan J https://orcid.org/0000-0003-0722-3058

Smolka, Marcus B https://orcid.org/0000-0001-9952-2885

Yu, Haiyuan https://orcid.org/0000-0001-7597-6049

Miller, Elizabeth A https://orcid.org/0000-0002-1033-8369

Fromme, J Christopher https://orcid.org/0000-0002-8837-0473

Funding for this research was provided by:

National Institute of General Medical Sciences (R01GM116942)

National Institute of General Medical Sciences (R01GM124559)

National Institute of General Medical Sciences (R01GM097272)

National Institute of General Medical Sciences (R35GM136258)

National Institute of General Medical Sciences (R01GM125639)

Division of Biological Infrastructure (DBI‐1661380)

Medical Research Council (MRC_UP_1201/10)

Division of Graduate Education (DGE‐1650441)

Alfred P. Sloan Foundation

Accepted Manuscript valid from 2021-05-21

Version of Record valid from 2021-05-21