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Maehara, Kazumitsu https://orcid.org/0000-0002-2933-5176

Tomimatsu, Kosuke https://orcid.org/0000-0003-3733-6686

Harada, Akihito

Tanaka, Kaori

Sato, Shoko

Fukuoka, Megumi

Okada, Seiji

Handa, Tetsuya https://orcid.org/0000-0001-7707-0195

Kurumizaka, Hitoshi https://orcid.org/0000-0001-7412-3722

Saitoh, Noriko

Kimura, Hiroshi https://orcid.org/0000-0003-0854-083X

Ohkawa, Yasuyuki https://orcid.org/0000-0001-6440-9954

Funding for this research was provided by:

Precursory Research for Embryonic Science and Technology (JPMJPR2026)

Precursory Research for Embryonic Science and Technology (JPMJPR19K7)

Japan Society for the Promotion of Science (JP19H05425)

Japan Society for the Promotion of Science (JP20H05368)

Japan Society for the Promotion of Science (JP18H05534)

Japan Society for the Promotion of Science (JP20H00449)

Japan Society for the Promotion of Science (JP18H04802)

Japan Society for the Promotion of Science (JP18H05527)

Japan Society for the Promotion of Science (JP19H05244)

Japan Society for the Promotion of Science (JP20K21398)

Japan Society for the Promotion of Science (JP20H00456)

Japan Society for the Promotion of Science (JP20H04846)

Japan Society for the Promotion of Science (JP18H05527)

Japan Society for the Promotion of Science (JP17H01417)

Japan Society for the Promotion of Science (JP19H04970)

Japan Society for the Promotion of Science (JP19H03158)

Japan Society for the Promotion of Science (JP20H05393)

Japan Society for the Promotion of Science (JP18K19432)

Japan Society for the Promotion of Science (JP19H03211)

Japan Society for the Promotion of Science (JP21H05755)

Japan Society for the Promotion of Science (JP21H00430)

Japan Society for the Promotion of Science (JP21H05292)

Japan Agency for Medical Research and Development (JP20ek0109489h0001)

Japan Agency for Medical Research and Development (JP19am0101076)

Japan Agency for Medical Research and Development (JP19am0101105)

Japan Agency for Medical Research and Development (JP20am0101076)

Core Research for Evolutional Science and Technology (JPMJCR16G1)

Exploratory Research for Advanced Technology (JPMJER1901)

Accepted Manuscript valid from 2021-11-03

Version of Record valid from 2021-11-03